Meanwhile, Khrone is at work on Ix and Dan (Caladan). The infiltration of Face Dancers in to the Ixian industry is widespread. The Face Dancers made it seem like the Ixians had figured out a way to navigate foldspace without a melange-addicted and mutated Guild Navigator. "Even the Chief Fabricator assumed that the recent breakthroughs were based on real Ixian knowledge and ingenuity, not brought in from the Outside Enemy." (Hunters of Dune, p. 295)
There is growing underlying suspicion, but no evidence and no desire by most to dig deeper. In a meeting with Uxtal of the Lost Tleilaxu, Hellica of the Honored Matres, and a Guild Navigator concerning spice production using axlotl tanks, the Navigator senses the paradox of the Lost Tleilaxu.
"If your people are so ignorant, how did they create Face Dancers so superior to any previous ones?" the Navigator asked. Uxtal shuddered, knowing -- now -- that his people had not, after all, created Khrone or his superior breed of shape-shifters. Apparently, they had merely been found out in the Scattering.
(Hunters of Dune, pp. 342 - 343)